low fiber, high sugar, very low calcium, A 1:2 calcium:phoshorous ratio (too high phosphorous)
Compared to turnip greens squash has 1/3 the fiber, 3x the sugar, 1/10 thae calcium and about the same phosphorous. Turnip greens have a calcium phosphorous ratio of 4.52:1
Russian tortoises are designed to eat high fiber, high calcium and low protein weeds, grasses and greens.